organic substance metabolic process organic substance metabolic process organic substance metabolic process organic substance metabolic process organic substance metabolic process organic substance metabolic process organic substance metabolic process organic substance metabolic process organic substance metabolic process organic substance metabolic process organic substance metabolic processorganic substance metabolic process organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound binding heterocyclic compound binding ion binding organic cyclic compound bindingorganic cyclic compound binding heterocyclic compound bindingheterocyclic compound binding ion bindingion binding cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cell organelle membrane-enclosed lumen cellcell organelleorganelle membrane-enclosed lumenmembrane-enclosed lumen