primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic process organic substance metabolic process catabolic process primary metabolic processprimary metabolic process organic substance metabolic processorganic substance metabolic process catabolic processcatabolic process hydrolase activity hydrolase activity hydrolase activity hydrolase activity hydrolase activity hydrolase activity hydrolase activity hydrolase activity hydrolase activity hydrolase activity hydrolase activityhydrolase activity cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cell organelle membrane-enclosed lumen extracellular region cellcell organelleorganelle membrane-enclosed lumenmembrane-enclosed lumen extracellular regionextracellular region