Manteia - Molecule gene

ENSMUSG00000028173

Mus musculus

Features

Gene ID:	ENSMUSG00000028173

Biological name :	Wls

Synonyms :	Q6DID7 / Wls / wntless WNT ligand secretion mediator

Possible biological names infered from orthology :	Q5T9L3

Species:	Mus musculus

Chr. number:	3
Strand:	1
Band:	H4
Gene start:	159839672
Gene end:	159938664

Corresponding Affymetrix probe sets:	10497149 (MoGene1.0st) 1423824_at (Mouse Genome 430 2.0 Array) 1423825_at (Mouse Genome 430 2.0 Array) 1437434_a_at (Mouse Genome 430 2.0 Array) 1458319_at (Mouse Genome 430 2.0 Array)

Cross references:	Ensembl peptide - ENSMUSP00000067898 Ensembl peptide - ENSMUSP00000142774 Ensembl peptide - ENSMUSP00000143475 NCBI entrez gene - 68151 See in Manteia. MGI - MGI:1915401 RefSeq - XM_006501964 RefSeq - NM_001356350 RefSeq - NM_026582 RefSeq Peptide - NP_001343279 RefSeq Peptide - NP_080858 swissprot - Q6DID7 swissprot - A0A0G2JEH4 Ensembl - ENSMUSG00000028173

See expression report in BioGPS
See gene description in Wikigenes
See co-cited genes in PubMed

Ortholog prediction (from Ensembl)

Ortholog name	ID	Species
wls	ENSDARG00000009534	Danio rerio
WLS	ENSGALG00000011238	Gallus gallus
WLS	ENSG00000116729	Homo sapiens

Paralog prediction (from Ensembl)

Paralog name

ID

Similarity(%)

No match

Protein motifs (from Interpro)

Interpro ID	Name
IPR009551	Protein wntless

Gene Ontology (GO)

Type	GO ID	Term	Ev.Code
biological_process	GO:0001707	mesoderm formation	IMP
biological_process	GO:0006886	intracellular protein transport	ISO
biological_process	GO:0007275	multicellular organism development	IEA
biological_process	GO:0009948	anterior/posterior axis specification	IMP
biological_process	GO:0016055	Wnt signaling pathway	IMP
biological_process	GO:0030177	positive regulation of Wnt signaling pathway	ISS
biological_process	GO:0030901	midbrain development	IMP
biological_process	GO:0030902	hindbrain development	IMP
biological_process	GO:0031017	exocrine pancreas development	IMP
biological_process	GO:0061355	Wnt protein secretion	ISO
biological_process	GO:0061357	positive regulation of Wnt protein secretion	ISS
biological_process	GO:0090263	positive regulation of canonical Wnt signaling pathway	IMP
cellular_component	GO:0000139	Golgi membrane	ISO
cellular_component	GO:0005768	endosome	IEA
cellular_component	GO:0005769	early endosome	ISO
cellular_component	GO:0005783	endoplasmic reticulum	IEA
cellular_component	GO:0005789	endoplasmic reticulum membrane	ISO
cellular_component	GO:0005794	Golgi apparatus	ISO
cellular_component	GO:0005802	trans-Golgi network	ISO
cellular_component	GO:0005886	plasma membrane	ISO
cellular_component	GO:0016020	membrane	IEA
cellular_component	GO:0016021	integral component of membrane	IEA
cellular_component	GO:0030659	cytoplasmic vesicle membrane	IEA
cellular_component	GO:0031410	cytoplasmic vesicle	ISO
cellular_component	GO:0031901	early endosome membrane	IEA
cellular_component	GO:0032590	dendrite membrane	IEA
cellular_component	GO:0032839	dendrite cytoplasm	IEA
molecular_function	GO:0005515	protein binding	IPI
molecular_function	GO:0017147	Wnt-protein binding	ISO
molecular_function	GO:0031852	mu-type opioid receptor binding	IEA

Pathways (from Reactome)

Pathway description

WNT ligand biogenesis and trafficking

Phenotype (from MGI, Zfin or HPO)

ID	Phenotype	Definition	Genetic BG
MP:0000060	delayed bone ossification	"late onset of the formation of bone" [J:40203]	Show Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0000074	abnormal neurocranium morphology	"malformed bones of the skull enclosing the brain" [Stedman s Medical Dictionary:ISBN 0-683-40008-8, J:55583]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0000111	cleft palate	"congenital fissure of the tissues normally uniting to form the palate" [Stedman s Medical Dictionary:ISBN 0-683-40008-8, J:49840]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0000219	increased neutrophil count	"greater than normal neutrophil numbers" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0000324	increased mast cell number	"greater than expected number of the granulated cells found in most tissues and which contain large amounts of histamine and heparin in granules" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Ctnnb1^tm2Kem/Ctnnb1^tm2Kem,Tg(Krt1-15-cre/PGR)22Cot/0 Genetic Background: involves: 129S1/Sv * 129X1/SvJ * C57BL/6 * SJL Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0000352	decreased cell proliferation	"less than the normal growth and reproduction of similar cells" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6 Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0000377	abnormal hair follicle	"malformed invagination of the epidermis from which the hair shaft develops" [J:65031]	Show Allelic Composition: Ctnnb1^tm2Kem/Ctnnb1^tm2Kem,Tg(Krt1-15-cre/PGR)22Cot/0 Genetic Background: involves: 129S1/Sv * 129X1/SvJ * C57BL/6 * SJL
MP:0000379	reduced hair follicle number	"fewer number of the epidermal invaginations from which the hair shaft develops" [J:15108]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0000416	sparse fur	"fur that is significantly less dense" [J:46854]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6 Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Tg(Foxn1-cre)1Tbo/0 Genetic Background: involves: C57BL/6
MP:0000428	abnormal craniofacial morphology	"anomalous structure or development of the face and/or cranium" [Stedman s Medical Dictionary:ISBN 0-683-40008-8]	Show Allelic Composition: Wls^tm1.2Whsu/Wls^tm1.2Whsu Genetic Background: involves: 129S6/SvEvTac
MP:0000547	short limbs	"reduced average length of the extremities" [MGI:CLS, J:61509]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6 Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0000550	abnormal forelimb morphology	"malformation of the entire anterior extremities" [MGI:tc]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0000556	abnormal hindlimb morphology	"malformation of the entire posterior extremities" [MGI:tc]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0000565	oligodactyly	"congenital condition in which some digits or parts of digits are missing" [Stedman s Medical Dictionary:ISBN 0-683-40008-8, J:53370]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0000572	abnormal autopod morphology	"malformation of the hand or foot" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6 Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0000613	abnormal salivary gland morphology	"malformed saliva-secreting glands of the oral cavity" [Stedman s Medical Dictionary , 27th edition:ISBN 0-683-40008-8, J:63764]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0000733	abnormal muscle development	"anomalous differentiation of muscle tissue" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0000847	abnormal metencephalon morphology	"any malformation or absence of the anterior part of the rhombencephalon" [Stedman s Medical Dictionary , 27th edition:ISBN 0-683-40008-8, MeSH:National Library of Medicine - Medical Subject Headings , 2003]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL
MP:0000850	absent cerebellum	"missing the part of the metencephalon that lies in the posterior cranial fossa dorsal to the pons and medulla that is concerned with the coordination of movement" [Stedman s Medical Dictionary:ISBN 0-683-40008-8, J:19212, MeSH:National Library of Medicine - Medical Subject Headings , 2003]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0001056	abnormal cranial nerve morphology	"any anomaly, deformity, or malformation of any of the twelve nerves that emerge from the cranium as opposed to the spinal nerves" [Stedman s Medical Dictionary:ISBN 0-683-40008-8, MGI:cls, J:45302]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0001190	reddish skin	"skin has a more intense ruddy appearance than normal, often due to capillary congestion" [J:65038]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0001192	scaly skin	"skin covered with shedding scales or flakes" [J:65146]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0001195	flaky skin	"shedding scales on the skin " [J:56641]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0001199	thin skin	"reduced thickness of the outer protective layer of the body" [Stedman s Medical Dictionary:ISBN 0-683-40008-8, J:49471]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0001216	abnormal epidermal layer morphology	"anomalous structure or development of the superficial epithelial portion of the skin" [Stedman s Medical Dictionary:ISBN 0-683-40008-8]	Show Allelic Composition: Ctnnb1^tm2Kem/Ctnnb1^tm2Kem,Tg(Krt1-15-cre/PGR)22Cot/0 Genetic Background: involves: 129S1/Sv * 129X1/SvJ * C57BL/6 * SJL
MP:0001243	abnormal dermal layer morphology	"malformed or atypical condition of the dermal layer of the skin" [Stedman s Medical Dictionary:ISBN 0-683-40008-8, J:56777]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0001244	thin dermal layer	"reduced thickness of the dermis" [J:65030]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0001265	reduced body size	"smaller than average body weight, height and/or length of an organism compared to controls" [Stedman s Medical Dictionary , 27th edition:ISBN 0-683-40008-8, J:23170]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0001428	adipsia	"absence of thirst or lack of desire to drink " [Stedman s Medical Dictionary:ISBN 0-683-40008-8, J:30404]	Show Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0001674	abnormal triploblastic development	"abnormal development of the three primary germ layers" [Stedman s Medical Dictionary , 27th edition:ISBN 0-683-40008-8, tc:Teresa Chu , Mouse Genome Informatics Curator]	Show Allelic Composition: Hsd11b1^tm1.1Ggla/Hsd11b1^tm1.1Ggla Genetic Background: involves: 129P2/OlaHsd
MP:0001675	abnormal ectoderm development	"failure or abnormality in the formation of the ectoderm during gastrulation" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Hsd11b1^tm1.1Ggla/Hsd11b1^tm1.1Ggla Genetic Background: involves: 129P2/OlaHsd
MP:0001676	abnormal apical ectodermal ridge	"malformation of the multilayered ectodermal region at the tip of a limb bud necessary for the proper development of the underlying mesenchyme" [MGI:tc]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0001679	thin apical ectodermal ridge	"decrease in the thickness of the multilayered ectodermal region at the tip of a limb bud necessary for the proper development of the underlying mesenchyme" [MGI:tc]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0001683	absent mesoderm	"missing or failure to differentiate the middle primary germ layer " [J:40594]	Show Allelic Composition: Hsd11b1^tm1.1Ggla/Hsd11b1^tm1.1Ggla Genetic Background: involves: 129P2/OlaHsd Allelic Composition: Wls^tm1.2Whsu/Wls^tm1.2Whsu Genetic Background: involves: 129S6/SvEvTac
MP:0001685	abnormal endoderm development	"failure or abnormality in the formation of the endoderm during gastrulation" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Hsd11b1^tm1.1Ggla/Hsd11b1^tm1.1Ggla Genetic Background: involves: 129P2/OlaHsd
MP:0001693	failure of primitive streak formation	"inability to form the epiblast ridge from which arises the germ layers of the embryo" [Stedman s Medical Dictionary:ISBN 0-683-40008-8, MGI:cls, J:33880]	Show Allelic Composition: Hsd11b1^tm1.1Ggla/Hsd11b1^tm1.1Ggla Genetic Background: involves: 129P2/OlaHsd Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,Tg(EIIa-cre)C5379Lmgd/0 Genetic Background: involves: 129S/SvEv * FVB/N * SJL Allelic Composition: Wls^tm1.2Whsu/Wls^tm1.2Whsu Genetic Background: involves: 129S6/SvEvTac
MP:0001730	embryonic growth arrest	"the cessation of development beyond a particular stage" [J:17509]	Show Allelic Composition: Kitl^Sl-10H/Kitl⁺ Genetic Background: involves: 101/H * C3H/HeH Allelic Composition: Wls^tm1.2Whsu/Wls^tm1.2Whsu Genetic Background: involves: 129S6/SvEvTac
MP:0001732	postnatal growth retardation	"slow or limited development after birth " [Stedman s Medical Dictionary:ISBN 0-683-40008-8]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0001823	thymus hypoplasia	"small size due to reduced cell number in the thymus" [J:23255]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0002073	abnormal hair growth	"absence or reduced amount of hair or abnormal onset of hair growth cycle or development" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Prdm4^{tm1b(EUCOMM)Liz}/Prdm4^{tm1b(EUCOMM)Liz} Genetic Background: involves: 129S/SvEv * C57BL/6 * CBA
MP:0002081	perinatal lethality	"death anytime between E18.5 and postnatal day 1 " [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,Tg(EIIa-cre)C5379Lmgd/0 Genetic Background: involves: 129S/SvEv * FVB/N * SJL
MP:0002083	premature death	"death after 3 weeks of age, but before normal life span" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0002084	abnormal developmental patterning	"abnormal systematic arrangement of the developing body along an axis" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Hsd11b1^tm1.1Ggla/Hsd11b1^tm1.1Ggla Genetic Background: involves: 129P2/OlaHsd
MP:0002092	abnormal eye morphology	"abnormal development of the eye tissues resulting in morphological abnormality or abnormal structure of the visual system" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0002100	abnormal tooth morphology	"atypical size, shape or hard tissue structure of the teeth " [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0002102	abnormal ear morphology	"structural or developmental anomaly of any of the structures involved in the ear or vestibular system" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Prdm4^{tm1b(EUCOMM)Liz}/Prdm4^{tm1b(EUCOMM)Liz} Genetic Background: involves: 129S/SvEv * C57BL/6 * CBA
MP:0002145	abnormal T lymphocyte development	"atypical production of or inability to produce mature T cells, and/or accumulation of T cell precursors" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0002151	abnormal neural tube morphology/development	"anomalous structure of or development of the embryonic neural tube resulting in structural anomaly" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL
MP:0002152	abnormal brain morphology	"anomalous structure of or abnormal development of the brain, one of the two components of the central nervous system and the center of thought and emotion; controls coordination, bodily activities and the interpretation of information from the senses (sight, hearing, smell, etc.)" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator, http://cancerweb.ncl.ac.uk]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0002233	abnormal nose morphology	"any structural anomaly of the organ that is specialized for smell and is part of the respiratory system " [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator, MeSH:National Library of Medicine - Medical Subject Headings , 2003]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0002444	abnormal T cell physiology	"anomalous response of T lymphocytes" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0002655	abnormal keratinocyte morphology	"structural or developmental anomaly of the cells of the epidermis that produce keratin in the process of differentiating into the dead and fully keratinized cells of the stratum corneum" [Stedman s Medical Dictionary:ISBN 0-683-40008-8]	Show Allelic Composition: Ctnnb1^tm2Kem/Ctnnb1^tm2Kem,Tg(Krt1-15-cre/PGR)22Cot/0 Genetic Background: involves: 129S1/Sv * 129X1/SvJ * C57BL/6 * SJL
MP:0002657	chondrodystrophy	"abnormal development of the cartilage primordia of long bones, resulting in abnormally short extremities but normal head and trunk" [il:Ira Lu , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0002764	short tibia	" reduced length of the medial and larger bone of the lower leg" [J:12736, pvb:Pierre Vanden Borre , Mouse Genome Informatics Curator]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0002765	short fibula	"reduced length of the lateral and smaller of bone of the lower leg" [pvb:Pierre Vanden Borre , Mouse Genome Informatics Curator, J:12736]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0002796	impaired skin barrier function	"impaired ability of the skin to regulate water loss; frequently leads to dehydration" [J:56641]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0003413	hair follicle degeneration	"a retrogressive impairment of function or destruction of the hair follicle" [monikat:Monika Tomczuk, Mouse Genome Informatics Curator, J:94517]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0003644	thymus atrophy	"wasting of the thymus resulting in reduced size " [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0003704	abnormal hair follicle development	"anomalous development of the epidermis from which the hair shaft develops " [monikat:Monika Tomczuk, Mouse Genome Informatics Curator]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0003854	abnormal forelimb stylopod morphology		Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0003857	abnormal hindlimb zeugopod morphology		Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0003864	abnormal midbrain development	"anomaly in the formation of or the patterning of the part of the brainstem developing from the middle of the three primary cerebral vesicles of the embryo" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator, Stedman s Medical Dictionary , 27th edition:ISBN 0-683-40008-8]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0003935	abnormal craniofacial development	"anomaly in the process of forming the face and/or cranium" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0003944	abnormal T cell subpopulation ratio	"deviation from the standard ratios of T cell subpopulations (e.g. double positive to single positive ratio) compared to control samples" [mberry:Melissa Berry, Genetics Resources Curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0004096	abnormal midbrain-hindbrain boundary development	"anomaly in the formation of the midbrain-hindbrain domain, comprised of the mesencephalic vesicle and the first rhombencephalic vesicle at early somitogenesis stages; an organizing center at the boundary patterns the midbrain and hindbrain primordia of the neural plate" [csmith:Cynthia L. Smith , Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0004126	thin hypodermis	"reduced thickness of the innermost and thickest layer of the skin composed of loose connective tissue containing adipoctyes" [anna:Anna Anagnostopoulos, Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0004947	skin inflammation	"local accumulation of fluid, plasma proteins, and leukocytes in the skin" [MGI:csmith "Cynthia L. Smith, Mouse Genome Informatics Curator"]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0004956	decreased thymus weight	"reduction in the average weight of the primary lymphoid organ that is required for maturation of T cells" [MGI:csmith "Cynthia L. Smith, Mouse Genome Informatics Curator"]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0005018	decreased T cell number	"fewer than normal T cell numbers" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0005090	increased double-negative T cells count	"greater than expected numbers of the subset of T cells found in the thymus that express neither CD4 nor CD8" [Immunobiology , The Immune System in Health and Disease:ISBN 0-8153-1691-7]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0005092	decreased double-positive T cell count	"less than the expected number of the subset of T cells found in the thymus that express both CD4 and CD8" [Immunobiology , The Immune System in Health and Disease:ISBN 0-8153-1691-7]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0005221	abnormal rostral-caudal axis patterning	"anomaly in the development or formation of the axis that runs from the head to the tail of the body" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Wls^tm1.2Whsu/Wls^tm1.2Whsu Genetic Background: involves: 129S6/SvEvTac
MP:0005274	abnormal viscerocranium morphology	"anomalous structure or formation of the part of the skull that comprises the facial bones " [Stedman s Medical Dictionary:ISBN 0-683-40008-8]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0005348	increased T cell proliferation	"increased clonal expansion of naive T cells in response to stimuli" [Immunobiology , The Immune System in Health and Disease:ISBN 0-8153-1691-7, cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0005425	increased macrophage count	"greater than the normal numbers of macrophages" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator, J:73418]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0005501	abnormal skin physiology	"atypical function of the membranous protective covering of the body" [cwg:Carroll-Ann W. Goldsmith , Mouse Genome Informatics curator, Stedman s Medical Dictionary:ISBN 0-683-40008-8]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0005503	abnormal tendon morphology	"structural anomaly of the fibrous bands or cords of connective tissue at the ends of muscle fibers that attach muscles to bones and other structures" [MeSH:National Library of Medicine - Medical Subject Headings , 2003, J:45509]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0005504	abnormal ligament morphology	"structural anomaly of the flexible bands of fibrous tissue that connect the articular extremities of bones " [MeSH:National Library of Medicine - Medical Subject Headings , 2003, J:45509]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0005508	abnormal skeleton morphology	"malformation of the bony framework of the body in vertebrates" [Stedman s Medical Dictionary:ISBN 0-683-40008-8]	Show Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0005666	abnormal adipose tissue physiology	"functional anomaly of the connective tissue composed of fat cells enmeshed in areolar tissue" [cwg:Carroll-Ann W. Goldsmith, Mouse Genome Informatics curator, MeSH:National Library of Medicine - Medical Subject Headings, 2003]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0006042	increased apoptosis	"greater than normal programmed cell death" [RGD:Rat Genome Database submission, smb:Susan M Bello, Mouse Genome Informatics Curator]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6 Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0006101	absent tegmentum	"absence or loss of the floor of the midbrain which extends from the substantia nigra to the level of the cerebral aqueduct" [smb:Susan M Bello, Mouse Genome Informatics Curator, anna:Anna Anagnostopoulos, Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0006106	absent tectum	"absence or loss of the rostral part of the midbrain roof" [anna:Anna Anagnostopoulos, Mouse Genome Informatics Curator, smb:Susan M Bello, Mouse Genome Informatics Curator]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0006108	abnormal hindbrain development	"anomaly in the formation or pattering of the caudal region of the brain" [smb:Susan M Bello, Mouse Genome Informatics Curator, J:93573]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0006279	abnormal limb development	"anomaly in the formation of the limbs" [MGI:smb "Susan M. Bello, Mouse Genome Informatics Curator"]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6 Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0006280	abnormal digit development	"anomaly in the formation of the digits" [J:70592, MGI:smb "Susan M. Bello, Mouse Genome Informatics Curator"]	Show Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0008052	abnormal serous gland morphology	"any structural anomaly of glands that secrete watery albuminous material that often contains enzymes" [ISBN:0-683-40008-8 "Stedman s Medical Dictionary, 27th edition"]	Show Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0008347	decreased gamma-delta T cell number	"reduced number of immature or mature T cells expressing an gamma-delta T cell receptor complex" [CL:0000798, ISBN:0781735149]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0008859	abnormal hair cycle catagen phase	"anomaly in the phase of the hair growth cycle during which cell proliferation ceases, the hair follicle shortens, and an anchored hair is produced" [MGI:llw2 "Linda Washburn, Mouse Genome Informatics Curator"]	Show Allelic Composition: Ctnnb1^tm2Kem/Ctnnb1^tm2Kem,Tg(Krt1-15-cre/PGR)22Cot/0 Genetic Background: involves: 129S1/Sv * 129X1/SvJ * C57BL/6 * SJL
MP:0008934	absent choroid plexus	"absence of the fringe of the tela choroidea of the brain ventricles" [ISBN:0-683-40008-8 "Stedman s Medical Dictionary, 27th edition"]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL Allelic Composition: Wls^tm1.1Whsu/Wls^tm1.1Whsu,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S6/SvEvTac * C57BL/6J * CBA/J
MP:0009111	pancreas hypoplasia	"underdevelopment or reduced size of the pancreas, usually due to a reduced number of cells" [MGI:csmith "Cynthia L. Smith, Mouse Genome Informatics Curator"]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL
MP:0009583	increased keratinocyte proliferation	"increase in the expansion rate of keratinocytes by cell division" [MGI:anna "Anna V. Anagnostopoulos, Mouse Genome Informatics Curator"]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0009795	epidermal spongiosis	"intercellular edema within the epidermis" [MGI:llw2 "Linda Washburn, Mouse Genome Informatics Curator"]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0009931	abnormal skin appearance	"skin that looks different from the usual state" [MGI:llw2 "Linda Washburn, Mouse Genome Informatics Curator"]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0010743	delayed suture closure	"late onset of the fusion of the bones of the skull" [MGI:smb "Susan M. Bello, Mouse Genome Informatics Curator"]	Show Allelic Composition: Rgs11^tm1Lex/Rgs11^tm1Lex,Rgs7^tm1Cjch/Rgs7^tm1Cjch Genetic Background: involves: 129S5/SvEvBrd * C57BL/6
MP:0011015	decreased body surface temperature	"reduced degree of heat natural to skin-environment interface of a living being" [MGI:smb "Susan M. Bello, Mouse Genome Informatics Curator"]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6
MP:0011084	partial lethality at weaning	"the appearance of lower than Mendelian ratios of organisms of a given genotype due to death of some, but not all of the organisms at weaning age" [MGI:csmith]	Show Allelic Composition: Wls^tm1Xzg/Wls^tm1Xzg,Tg(Prrx1-cre)1Cjt/0 Genetic Background: involves: C57BL/6J * SJL/J
MP:0011092	complete embryonic lethality	"death of all organisms of a given genotype in a population within the embryonic period prior to organogenesis (Mus: prior to E14)" [MGI:csmith]	Show Allelic Composition: Gt(ROSA)26Sor^{tm1.1(Wls/YFP)Mbtr}/Gt(ROSA)26Sor⁺ Genetic Background: involves: C57BL/6
MP:0011098	complete embryonic lethality during organogenesis	"death of all organisms of a given genotype in a population between embryo turning and the completion of organogenesis (Mus: E9 to less than E14)" [MGI:csmith]	Show Allelic Composition: Hsd11b1^tm1.1Ggla/Hsd11b1^tm1.1Ggla Genetic Background: involves: 129P2/OlaHsd Allelic Composition: Wls^tm1.2Whsu/Wls^tm1.2Whsu Genetic Background: involves: 129S6/SvEvTac
MP:0011106	partial embryonic lethality before somite formation	"the appearance of lower than Mendelian ratios of organisms of a given genotype due to death of some, but not all of the organisms between the point of implantation and somite formation (Mus: E4.5 to less than E8)" [MGI:csmith]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,Tg(EIIa-cre)C5379Lmgd/0 Genetic Background: involves: 129S/SvEv * FVB/N * SJL
MP:0012087	absent midbrain	"absence of the brain region derived from the middle of the three cerebral vesicles of the embryo; this region controls sensory and motor functions in the adult, including eye movement and coordination of auditory and visual reflexes" [MGI:anna]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL
MP:0012133	absent midbrain-hindbrain boundary	"absence of the midbrain-hindbrain domain of the embryonic brain that is comprised of the mesencephalic vesicle and the first rhombencephalic vesicle at early somitogenesis stages; normally, an organizing center located at the boundary patterns the midbrain and hindbrain primordia of the neural plate" [MGI:anna]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL
MP:0012138	decreased forebrain size	"size reduction or truncation of the anterior of the three primary divisions of the developing chordate brain or the corresponding part of the adult brain (in vertebrates, includes especially the cerebral hemispheres, the thalamus, and the hypothalamus and especially in higher vertebrates is the main control center for sensory and associative information processing, visceral functions, and voluntary motor functions)" [GO:0048853]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL
MP:0012173	short rostral-caudal axis	"length reduction or truncation of the axis that runs from the head to the tail of the body" [MGI:anna]	Show Allelic Composition: Wls^tm1.1Lan/Wls^tm1.1Lan,H2afv^{Tg(Wnt1-cre)11Rth}/0 Genetic Background: involves: 129S/SvEv * C57BL/6J * CBA/J * SJL
MP:0013588	small thymus medulla	"decreased size of the inner area of thymus lobules" [MGI:Anna]	Show Allelic Composition: Wls^tm1.1Arte/Wls^tm1.1Arte,Krt14^tm1(cre)Wbm/Krt14⁺ Genetic Background: involves: 129P2/OlaHsd * C57BL/6

Interacting proteins (from Reactome)

Interactor ID	Name	Interaction type
ENSMUSG00000029462	Vps29 / Q9QZ88 / VPS29 retromer complex component / Q9UBQ0*	/ reaction / complex
ENSMUSG00000019804	Snx3 / sorting nexin 3 / O60493*	/ complex / reaction
ENSMUSG00000031696	Vps35 / Q9EQH3 / VPS35 retromer complex component / Q96QK1*	/ reaction / complex
ENSMUSG00000020078	P40336 / Vps26a / VPS26 retromer complex component A / O75436*	/ reaction / complex

1 s.

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